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tromere. Crossing over can greatly increase the genetic variability in gametes by associating alleles that were not previously joined. (We examine the molecular mechanism of this process in chapter 12.) Before crossing over takes place, densely staining nodules are visible, rst in zygonema and lasting through pachynema. These are called recombination nodules ( g. 3.22a); they are correlated with crossing over and presumably represent the enzymatic machinery present on the chromosomes. As the chromosomes shorten and thicken further in diplonema, each chromosome can be seen to be made of two sister chromatids. Now the chromosome gures are referred to as tetrads because each is made up of four chromatids (see g. 3.19). At about this time, the synaptonemal complex disintegrates in all but the areas of the chiasmata (singular: chiasma), the X-shaped con gurations marking the places of crossing over ( g. 3.22b).Virtually all tetrads exhibit chiasmata; in cases in which no crossing over occurs, the tetrads tend to fall apart and segregate randomly. Thus, crossing over not only increases genetic diversity but also ensures the proper separation of homologous chromosomes. A meiosis-speci c form of cohesin keeps sister chromatids together. During the diplotene stage, chromosomes can again uncondense and become active. This is especially obvious in amphibians and birds, which produce a great amount of cytoplasmic nutrient for the future zygote. Recondensation of the chromosomes takes place at the end of diplonema. This stage can be very long; in human females, it begins in the fetus and does not complete until the egg is shed during ovulation, sometimes more than fty years later. As prophase I moves into diakinesis, the chromosomes become very condensed (see g. 3.19).

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Sampling rate Model number AT-MIO-16(L/H)-9 AT-MIO-16(L/H)-15 AT-MIO-16(L/H)-25 Typical case 100 ksamples/s 71 ksamples/s 45 ksamples/s Worst case 91 ksamples/s 59 ksamples/s 37 ksamples/s

spermatocytes of the pigeon, Columba livia. (Bar 1 m.) (b) A tetrad from the grasshopper, Chorthippus parallelus, at diplonema with ve chiasmata. ([a] From M. I. Pigozzi and

The timing of multiple A/D conversion is controlled either by the onboard counter/timer or by external timing signals The onboard sample rate clock and sample counter control the onboard A/D timing The AT-MIO-16 can generate both interrupts and DMA (direct memory access) requests on the PC/AT I/O channel The interrupt can be generated when 1 2 3 4 An A/D conversion is available to be read from the A/D buffer The sample counter reaches its terminal count An error occurs One of the onboard timer clocks generates a pulse

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On the other hand, DMA requests can be generated whenever an A/D measurement is available from the A/D buffer Analog Output: The AT-MIO-16 has two double-buffered multiplying 12-bit DACs that are connected to two analog output channels The resolution of the 12-bit DACs is 244 mV in the unipolar mode or 488 mV in the bipolar mode with the onboard 10-V reference Finer resolutions can be achieved by using smaller voltages on the external reference The analog output channels have an accuracy of 05 LSB and a differential linearity of 1 LSB Voltage offset and gain error can be trimmed to zero Digital I/O: The AT-MIO-16 has eight digital I/O lines that are divided into two four-bit ports The digital input circuitry has an eight-bit register that continuously reads the eight digital I/O lines, thus making read-back capability possible for the digital output ports, as well as reading incoming signals The digital I/O lines are TTL-compatible Counter/Timer: The AT-MIO-16 uses the AM9513A counter/timer for time-related functions The AM9513A contains ve independent 16-bit counter/timers

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A. J. Solari, Recombination Nodule Mapping and Chiasma Distribution in Spermatocytes of the Pigeon, Columba livia, in Genome, 42: 308 314, 1999. Reprinted by permission. [b] Courtesy of Bernard John.)

R v(t) = 1414 sin ( t) + ~ ( = 377 rad/s) _

There are several ways to determine whether a given population conforms to the Hardy-Weinberg equilibrium at a particular locus. However, the question usually arises when there is just a single sample from a population, representing only one generation. Can the existence of the Hardy-Weinberg equilibrium be determined with just one sample The answer is that we can determine whether the three genotypes (AA, Aa, and aa) occur with the frequencies p2, 2pq, and q2. If they do, then the population is considered to be in Hardy-Weinberg proportions; if not, then the population is not considered to be in Hardy-Weinberg proportions.

Known Quantities: Source voltage and frequency, load resistance and inductance values Find: Pav and p(t) for the RL load Figure 75

i(t)

To determine whether observed and expected allelic frequencies are the same, we can use the chi-square statistical test. In a chi-square test, we compare an observed number with an expected number. In this case, the observed values are the actual numbers of the three genotypes in the sample, and the expected values come from the prediction that the genotypes will occur in the p2, 2pq, and q2 proportions. An analysis for the Ohio MN blood-type data is presented in table 19.2.The agreement between observed and expected numbers is very good, obvious even before the calculation of the chi-square value. Since the critical chi-square for one degree of freedom at the 0.05 level is 3.841 (see table 4.4), we nd that

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Schematics, Diagrams, Circuits, and Given Data: v(t) = 1414 sin(377t) V; R = 4 Assumptions: Use rms values for all phasor quantities in the problem

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